Project 3932: E. Vlachos, G. L. Georgalis, S. Roussiakis, M. Böhme, G. Theodorou. 2019. The Pikermian tortoises (Testudines, Testudinidae) from the late Miocene of the South Balkans. Journal of Vertebrate Paleontology. 39 (5):e1711520.
Specimen: † Hesperotestudo osborniana (unvouchered)
View: ventral

Abstract

This article deals with the fossil tortoises of one of the most iconic fossil localities of the Neogene of the Old World, the upper Miocene locality of Pikermi, near Athens, Greece. We describe the type, previously published, and new material of the fossil tortoises from Pikermi, along with new material from the coeval Azmaka 6 locality in Bulgaria. This combined information results in the complete revision of the ‘marble tortoise’ Testudo marmorum, which is the first turtle species ever named from Greece. Besides the challenges and difficulties of working with material from the old collections of Pikermi, we present a complete revision of this species and identify its presence outside its type locality in the South Balkan Peninsula. The presence of numerous shells of the ‘marble tortoise’ allows us to discuss the observed variation and attempt to interpret it as the result of intraspecific variation, ontogenetic changes, or sexual dimorphism. For the first time, we observe the absence of the characteristic hypo-xiphiplastral hinge in female individuals of Testudo marmorum, which in turn challenges the traditional phylogenetic position of this species. We further describe previously published and new material of the giant tortoise Titanochelon from Pikermi. Based on the updated information, we can suggest that the Pikermian giant tortoise probably represents a distinct taxon. Our results signal Pikermi as one of the most important localities to understand the evolution of tortoises in the eastern Mediterranean during the last parts of the Neogene.


Read the article »

Article DOI: 10.1080/02724634.2019.1711520

Project DOI: 10.7934/P3932, http://dx.doi.org/10.7934/P3932
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 72 Taxa
  • 1 Specimen
  • 170 Characters
Total size of project's media files: 922.89k

Download Project SDD File
Total scored cells: 7942
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 170
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 397
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:0/170
Currently Viewing:
MorphoBank Project 3932
  • Creation Date:
    17 February 2021
  • Publication Date:
    17 February 2021
  • Project views: 4511

    Authors' Institutions

    • Museo Paleontológico Egidio Feruglio

    • Aristotle University of Thessaloniki

    • University of Athens

    • Università degli Studi di Torino (University of Turin)

    • Universitaet Tuebingen (Eberhard-Karls-Universitaet)

    • Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET)

    • Comenius University



    Members

    member name taxa specimens media chars character
    media
    labels
    cell scorings
    (scored, NPA, "-")
    cell
    medialabels
    rules
    MorphoBank Curator
    Project Administrator
    7211170007942
    (7660, 0, 282)
    000
    Evangelos Vlachos
    Full membership
    0000000
    (0, 0, 0)
    000


    Taxonomic Overview for Matrix 'M27236' (72 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Pleurodira
    Last Modified in 02/17/21
    23148126021002
    [2] Chelydra serpentina
    Last Modified in 02/17/21
    5150148017002
    [3] Kinosternon odoratus
    Last Modified in 02/17/21
    3155155012002
    [4] Caretta caretta
    Last Modified in 02/17/21
    728888010002
    [5] Apalone spinifera
    Last Modified in 02/17/21
    279494049002
    [6] Platysternon megacephalum
    Last Modified in 02/17/21
    1514814807002
    [7] Batagur baska
    Last Modified in 02/17/21
    416416204002
    [8] Mauremys caspica
    Last Modified in 02/17/21
    716115805002
    [9] Siebenrockiella crassicollis
    Last Modified in 02/17/21
    1615014806002
    [10] Mauremys sinensis
    Last Modified in 02/17/21
    4012812604002
    [11] Chrysemys picta
    Last Modified in 02/17/21
    116516207002
    [12] Emys orbicularis
    Last Modified in 02/17/21
    3113413108002
    [13] Trachemys scripta
    Last Modified in 02/17/21
    816015804002
    [14] Gopherus agassizii
    Last Modified in 02/17/21
    816415903002
    [15] Chelonoidis carbonaria
    Last Modified in 02/17/21
    018816703002
    [16] Chelonoidis chilensis
    Last Modified in 02/17/21
    418316303002
    [17] Chelonoidis denticulata
    Last Modified in 02/17/21
    019616703002
    [18] Geochelone elegans
    Last Modified in 02/17/21
    5311711403002
    [19] Indotestudo elongata
    Last Modified in 02/17/21
    2614514202002
    [20] Manouria emys
    Last Modified in 02/17/21
    701019901002
    [21] Kinixys erosa
    Last Modified in 02/17/21
    12148147011002
    [22] Indotestudo forstenii
    Last Modified in 02/17/21
    316516403002
    [23] Testudo graeca
    Last Modified in 02/17/21
    217916404002
    [24] Chersine hermanni
    Last Modified in 02/17/21
    616816202002
    [25] Kinixys homeana
    Last Modified in 02/17/21
    2913413308002
    [26] Agrionemys horsfieldii
    Last Modified in 02/17/21
    5811611002002
    [27] Manouria impressa
    Last Modified in 02/17/21
    2913813803002
    [28] Testudo kleinmanni
    Last Modified in 02/17/21
    3113913405002
    [29] Testudo marginata
    Last Modified in 02/17/21
    79978902002
    [30] Chelonoidis nigra
    Last Modified in 02/17/21
    217316503002
    [31] Psammobates oculifera
    Last Modified in 02/17/21
    73979601002
    [32] Stigmochelys pardalis
    Last Modified in 02/17/21
    1315715403002
    [33] Gopherus polyphemus
    Last Modified in 02/17/21
    616516103002
    [34] Astrochelys radiata
    Last Modified in 02/17/21
    916015902002
    [35] Centrochelys sulcata
    Last Modified in 02/17/21
    217116602002
    [36] Malacochersus tornieri
    Last Modified in 02/17/21
    2152150018002
    [37] Gigantochersina ammon
    Last Modified in 02/17/21
    113635700002
    [38] † Testudo antiqua
    Last Modified in 02/17/21
    106666301002
    [39] Megalochelys atlas
    Last Modified in 02/17/21
    133413700002
    [40] † Titanochelon bacharidisi
    Last Modified in 02/17/21
    701029802002
    [41] † Hesperotestudo bermudae
    Last Modified in 02/17/21
    100737000002
    [42] † Titanochelon bolivari
    Last Modified in 02/17/21
    3513713203002
    [43] Ergilemys bruneti
    Last Modified in 02/17/21
    111605900002
    [44] † Paleotestudo canetotiana
    Last Modified in 02/17/21
    112585701002
    [45] † Achilemys cassouleti
    Last Modified in 02/17/21
    103676700002
    [46] Hadrianus castrensis
    Last Modified in 02/17/21
    113565601002
    [47] Hadrianus corsoni
    Last Modified in 02/17/21
    104666501002
    [48] Geochelone costaricensis
    Last Modified in 02/17/21
    112585800002
    [49] † Hesperotestudo crassiscutata
    Last Modified in 02/17/21
    107636300002
    [50] Testudo eocaenica
    Last Modified in 02/17/21
    114635600002
    [51] Taraschelon gigas
    Last Modified in 02/17/21
    125424203002
    [52] Chelonoidis gringorum
    Last Modified in 02/17/21
    130383802002
    [53] Chelonoidis hesterna
    Last Modified in 02/17/21
    108606002002
    [54] Ergilemys insolitus
    Last Modified in 02/17/21
    123524700002
    [55] Testudo kaiseni
    Last Modified in 02/17/21
    127424201002
    [56] Oligopherus laticuneus
    Last Modified in 02/17/21
    91787801002
    [57] † Testudo lunellensis
    Last Modified in 02/17/21
    111585801002
    [58] Cheirogaster maurini
    Last Modified in 02/17/21
    116515103002
    [59] Namibchersus namaquensis
    Last Modified in 02/17/21
    108626200002
    [60] Stylemys nebrascensis
    Last Modified in 02/17/21
    5511611401002
    [61] † Mesochersus orangeus
    Last Modified in 02/17/21
    135373401002
    [62] † Hesperotestudo osborniana
    Last Modified in 02/17/21
    6011211000002
    [63] † Testudo oughlamensis
    Last Modified in 02/17/21
    119535001002
    [64] Impregnochelys pachytestis
    Last Modified in 02/17/21
    130404000002
    [65] † Titanochelon perpiniana
    Last Modified in 02/17/21
    87807904002
    [66] Testudo promarginata
    Last Modified in 02/17/21
    111595900002
    [67] Testudo sharanensis
    Last Modified in 02/17/21
    115545401002
    [68] † Testudo shensiensis
    Last Modified in 02/17/21
    102686701002
    [69] Anhuichelys siaoshihensis
    Last Modified in 02/17/21
    105616104002
    [70] Pelorochelon soriana
    Last Modified in 02/17/21
    106666400002
    [71] † Titanochelon vitodurana
    Last Modified in 02/17/21
    81888702002
    [72] † Testudo marmorum
    Last Modified in 02/17/21
    105726401002


    Project views

    type number of views Individual items viewed (where applicable)
    Total project views4511
    Project overview351
    Matrix views273Data matrix (33 views); Matrix landing page (240 views);
    Media views897Media search (767 views); M739118 (130 views);
    Taxon list1909
    Bibliography178
    Views for media list461
    Specimen list442




    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project79
    Project downloads79