Project 3936: E. Vlachos, A. Pérez-García, S. Roussiakis, G. L. Georgalis, B. P. Kear. 2019. Late Miocene tortoises from Samos, Greece: implications for European Neogene testudinid systematics and distributions. Journal of Vertebrate Paleontology. 39 (6):e1722950.
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Geochelone elegans
whole organism