Project 3936: E. Vlachos, A. Pérez-García, S. Roussiakis, G. L. Georgalis, B. P. Kear. 2019. Late Miocene tortoises from Samos, Greece: implications for European Neogene testudinid systematics and distributions. Journal of Vertebrate Paleontology. 39 (6):e1722950.
Abstract
The world-famous upper Miocene fossil localities on the Aegean island of Samos in Greece have produced a rich fossil record that sheds light on the evolution of eastern Mediterranean terrestrial faunas over a one-million-year interval of the late Neogene. Fossils have been discovered on Samos since antiquity, although a succession of paleontological and commercial collecting expeditions over the last 130 years has resulted in specimens now being distributed throughout museums all over the world. Here, we survey the fossil tortoise remains from Samos, which are significant because they include early antecedents of the modern Testudo lineage, together with spectacular examples of the European Neogene gigantic testudinid †Titanochelon, which represents one of the largest-bodied terrestrial turtle taxa documented to date. All of the Samos fossils derive from the Mytilinii Formation, which spans the late MN11–early MN13 Neogene land mammal zones. The small-bodied tortoise remains include two incomplete shells that are morphologically consistent with basal testudonans and phylogenetically distinct from the coeval species Testudo marmorum found on mainland Greece. The Samos gigantic tortoise †‘Testudo’ schafferi was based on a spectacularly large skull and femur. However, we describe new plastron fragments, limb elements, and osteoderms that are compatible with †Titanochelon specimens from southern Greece and Anatolia. This could imply faunal links with the distinctive ‘Pikermian’ local assemblages from Asia Minor and concurs with the proposed late Miocene–Pliocene biogeographic segregation of large mammals from the eastern Aegean margin and Turkey relative to those occurring in northwestern Greece and the Balkan Peninsula.Read the article »
Article DOI: 10.1080/02724634.2019.1722950
Project DOI: 10.7934/P3936, http://dx.doi.org/10.7934/P3936
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 5227 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 170 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 395 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:145/25 |
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MorphoBank Project 3936
MorphoBank Project 3936
- Creation Date:
19 February 2021 - Publication Date:
19 February 2021 - Project views: 8506
- Matrix downloads: 8
Authors' Institutions
- Museo Paleontológico Egidio Feruglio
- Aristotle University of Thessaloniki
- University of Athens
- Università degli Studi di Torino (University of Turin)
- Uppsala University
- Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET)
- Universidad Nacional de Educación a Distancia (UNED), Madrid
- Comenius University
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
MorphoBank Curator Project Administrator | 42 | 1 | 1 | 170 | 0 | 0 | 5227 (5087, 0, 140) | 0 | 0 | 0 | ||||
Evangelos Vlachos Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M27242' (42 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Pleurodira Last Modified in 02/19/21 | 23 | 148 | 126 | 0 | 21 | 0 | 0 | 2 |
[2] Mauremys caspica Last Modified in 02/19/21 | 7 | 161 | 157 | 0 | 5 | 0 | 0 | 2 |
[3] Trachemys scripta Last Modified in 02/19/21 | 8 | 160 | 158 | 0 | 4 | 0 | 0 | 2 |
[4] Gopherus agassizii Last Modified in 02/19/21 | 8 | 164 | 159 | 0 | 3 | 0 | 0 | 2 |
[5] Chelonoidis carbonaria Last Modified in 02/19/21 | 0 | 188 | 155 | 0 | 3 | 0 | 0 | 2 |
[6] Chelonoidis chilensis Last Modified in 02/19/21 | 4 | 183 | 163 | 0 | 3 | 0 | 0 | 2 |
[7] Chelonoidis denticulata Last Modified in 02/19/21 | 0 | 196 | 166 | 0 | 3 | 0 | 0 | 2 |
[8] Geochelone elegans Last Modified in 02/19/21 | 53 | 117 | 114 | 0 | 3 | 0 | 0 | 2 |
[9] Indotestudo elongata Last Modified in 02/19/21 | 26 | 145 | 142 | 0 | 2 | 0 | 0 | 2 |
[10] Manouria emys Last Modified in 02/19/21 | 70 | 101 | 98 | 0 | 1 | 0 | 0 | 2 |
[11] Kinixys erosa Last Modified in 02/19/21 | 12 | 148 | 140 | 0 | 11 | 0 | 0 | 2 |
[12] Indotestudo forstenii Last Modified in 02/19/21 | 3 | 165 | 163 | 0 | 3 | 0 | 0 | 2 |
[13] Testudo graeca Last Modified in 02/19/21 | 2 | 179 | 163 | 0 | 4 | 0 | 0 | 2 |
[14] Chersine hermanni Last Modified in 02/19/21 | 6 | 168 | 161 | 0 | 2 | 0 | 0 | 2 |
[15] Kinixys homeana Last Modified in 02/19/21 | 29 | 134 | 133 | 0 | 8 | 0 | 0 | 2 |
[16] Agrionemys horsfieldii Last Modified in 02/19/21 | 58 | 116 | 110 | 0 | 2 | 0 | 0 | 2 |
[17] Manouria impressa Last Modified in 02/19/21 | 29 | 138 | 138 | 0 | 3 | 0 | 0 | 2 |
[18] Testudo kleinmanni Last Modified in 02/19/21 | 31 | 139 | 133 | 0 | 5 | 0 | 0 | 2 |
[19] Testudo marginata Last Modified in 02/19/21 | 79 | 97 | 89 | 0 | 2 | 0 | 0 | 2 |
[20] Chelonoidis nigra Last Modified in 02/19/21 | 2 | 173 | 165 | 0 | 3 | 0 | 0 | 2 |
[21] Psammobates oculifera Last Modified in 02/19/21 | 73 | 97 | 96 | 0 | 1 | 0 | 0 | 2 |
[22] Stigmochelys pardalis Last Modified in 02/19/21 | 13 | 157 | 154 | 0 | 3 | 0 | 0 | 2 |
[23] Gopherus polyphemus Last Modified in 02/19/21 | 6 | 165 | 161 | 0 | 3 | 0 | 0 | 2 |
[24] Astrochelys radiata Last Modified in 02/19/21 | 9 | 160 | 159 | 0 | 2 | 0 | 0 | 2 |
[25] Centrochelys sulcata Last Modified in 02/19/21 | 2 | 171 | 165 | 0 | 2 | 0 | 0 | 2 |
[26] Malacochersus tornieri Last Modified in 02/19/21 | 2 | 152 | 150 | 0 | 18 | 0 | 0 | 2 |
[27] † Testudo antiqua Last Modified in 02/19/21 | 106 | 67 | 62 | 0 | 1 | 0 | 0 | 2 |
[28] † Titanochelon bacharidisi Last Modified in 02/19/21 | 70 | 102 | 97 | 0 | 2 | 0 | 0 | 2 |
[29] † Titanochelon bolivari Last Modified in 02/19/21 | 35 | 137 | 132 | 0 | 3 | 0 | 0 | 2 |
[30] † Paleotestudo canetotiana Last Modified in 02/19/21 | 108 | 65 | 61 | 0 | 1 | 0 | 0 | 2 |
[31] † Achilemys cassouleti Last Modified in 02/19/21 | 103 | 67 | 65 | 0 | 0 | 0 | 0 | 2 |
[32] † Hadrianus corsoni Last Modified in 02/19/21 | 104 | 66 | 63 | 0 | 1 | 0 | 0 | 2 |
[33] † Testudo eocaenica Last Modified in 02/19/21 | 113 | 65 | 57 | 0 | 0 | 0 | 0 | 2 |
[34] Testudo kaiseni Last Modified in 02/19/21 | 127 | 42 | 42 | 0 | 1 | 0 | 0 | 2 |
[35] † Oligopherus laticuneus Last Modified in 02/19/21 | 91 | 78 | 78 | 0 | 1 | 0 | 0 | 2 |
[36] † Testudo marmorum Last Modified in 02/19/21 | 111 | 58 | 58 | 0 | 1 | 0 | 0 | 2 |
[37] † Titanochelon perpiniana Last Modified in 02/19/21 | 87 | 80 | 78 | 0 | 4 | 0 | 0 | 2 |
[38] Testudo promarginata Last Modified in 02/19/21 | 111 | 59 | 59 | 0 | 0 | 0 | 0 | 2 |
[39] † Testudo sharanensis Last Modified in 02/19/21 | 115 | 54 | 53 | 0 | 1 | 0 | 0 | 2 |
[40] † Pelorochelon soriana Last Modified in 02/19/21 | 106 | 70 | 64 | 0 | 0 | 0 | 0 | 2 |
[41] † Titanochelon vitodurana Last Modified in 02/19/21 | 81 | 88 | 87 | 0 | 2 | 0 | 0 | 2 |
[42] † Titanochelon schafferi Last Modified in 02/19/21 | 101 | 67 | 67 | 0 | 2 | 0 | 0 | 2 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 8506 | |
Project overview | 844 | |
Matrix views | 553 | Data matrix (64 views); Matrix landing page (489 views); |
Media views | 1859 | Media search (1566 views); M739129 (293 views); |
Taxon list | 3406 | |
Bibliography | 353 | |
Views for media list | 524 | |
Specimen list | 962 | |
Documents list | 5 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 85 | |
Matrix downloads | 8 | Data matrix (8 downloads); |
Project downloads | 77 |