Project 3936: E. Vlachos, A. Pérez-García, S. Roussiakis, G. L. Georgalis, B. P. Kear. 2019. Late Miocene tortoises from Samos, Greece: implications for European Neogene testudinid systematics and distributions. Journal of Vertebrate Paleontology. 39 (6):e1722950.
Specimen: Geochelone elegans (unvouchered)
View: whole organism

Abstract

The world-famous upper Miocene fossil localities on the Aegean island of Samos in Greece have produced a rich fossil record that sheds light on the evolution of eastern Mediterranean terrestrial faunas over a one-million-year interval of the late Neogene. Fossils have been discovered on Samos since antiquity, although a succession of paleontological and commercial collecting expeditions over the last 130 years has resulted in specimens now being distributed throughout museums all over the world. Here, we survey the fossil tortoise remains from Samos, which are significant because they include early antecedents of the modern Testudo lineage, together with spectacular examples of the European Neogene gigantic testudinid †Titanochelon, which represents one of the largest-bodied terrestrial turtle taxa documented to date. All of the Samos fossils derive from the Mytilinii Formation, which spans the late MN11–early MN13 Neogene land mammal zones. The small-bodied tortoise remains include two incomplete shells that are morphologically consistent with basal testudonans and phylogenetically distinct from the coeval species Testudo marmorum found on mainland Greece. The Samos gigantic tortoise †‘Testudo’ schafferi was based on a spectacularly large skull and femur. However, we describe new plastron fragments, limb elements, and osteoderms that are compatible with †Titanochelon specimens from southern Greece and Anatolia. This could imply faunal links with the distinctive ‘Pikermian’ local assemblages from Asia Minor and concurs with the proposed late Miocene–Pliocene biogeographic segregation of large mammals from the eastern Aegean margin and Turkey relative to those occurring in northwestern Greece and the Balkan Peninsula.


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Article DOI: 10.1080/02724634.2019.1722950

Project DOI: 10.7934/P3936, http://dx.doi.org/10.7934/P3936
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 42 Taxa
  • 1 Specimen
  • 170 Characters
Total size of project's media files: 226.23k

Download Project SDD File
Total scored cells: 5227
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 170
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 395
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:145/25
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MorphoBank Project 3936

    Authors' Institutions

    • Museo Paleontológico Egidio Feruglio

    • Aristotle University of Thessaloniki

    • University of Athens

    • Università degli Studi di Torino (University of Turin)

    • Uppsala University

    • Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET)

    • Universidad Nacional de Educación a Distancia (UNED), Madrid

    • Comenius University



    Members

    member name taxa specimens media chars character
    media
    labels
    cell scorings
    (scored, NPA, "-")
    cell
    medialabels
    rules
    MorphoBank Curator
    Project Administrator
    4211170005227
    (5087, 0, 140)
    000
    Evangelos Vlachos
    Full membership
    0000000
    (0, 0, 0)
    000


    Taxonomic Overview for Matrix 'M27242' (42 Taxa)

    taxon unscored
    cells
    scored
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    "-" cellscell images labels on
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    [1] Pleurodira
    Last Modified in 02/19/21
    23148126021002
    [2] Mauremys caspica
    Last Modified in 02/19/21
    716115705002
    [3] Trachemys scripta
    Last Modified in 02/19/21
    816015804002
    [4] Gopherus agassizii
    Last Modified in 02/19/21
    816415903002
    [5] Chelonoidis carbonaria
    Last Modified in 02/19/21
    018815503002
    [6] Chelonoidis chilensis
    Last Modified in 02/19/21
    418316303002
    [7] Chelonoidis denticulata
    Last Modified in 02/19/21
    019616603002
    [8] Geochelone elegans
    Last Modified in 02/19/21
    5311711403002
    [9] Indotestudo elongata
    Last Modified in 02/19/21
    2614514202002
    [10] Manouria emys
    Last Modified in 02/19/21
    701019801002
    [11] Kinixys erosa
    Last Modified in 02/19/21
    12148140011002
    [12] Indotestudo forstenii
    Last Modified in 02/19/21
    316516303002
    [13] Testudo graeca
    Last Modified in 02/19/21
    217916304002
    [14] Chersine hermanni
    Last Modified in 02/19/21
    616816102002
    [15] Kinixys homeana
    Last Modified in 02/19/21
    2913413308002
    [16] Agrionemys horsfieldii
    Last Modified in 02/19/21
    5811611002002
    [17] Manouria impressa
    Last Modified in 02/19/21
    2913813803002
    [18] Testudo kleinmanni
    Last Modified in 02/19/21
    3113913305002
    [19] Testudo marginata
    Last Modified in 02/19/21
    79978902002
    [20] Chelonoidis nigra
    Last Modified in 02/19/21
    217316503002
    [21] Psammobates oculifera
    Last Modified in 02/19/21
    73979601002
    [22] Stigmochelys pardalis
    Last Modified in 02/19/21
    1315715403002
    [23] Gopherus polyphemus
    Last Modified in 02/19/21
    616516103002
    [24] Astrochelys radiata
    Last Modified in 02/19/21
    916015902002
    [25] Centrochelys sulcata
    Last Modified in 02/19/21
    217116502002
    [26] Malacochersus tornieri
    Last Modified in 02/19/21
    2152150018002
    [27] † Testudo antiqua
    Last Modified in 02/19/21
    106676201002
    [28] † Titanochelon bacharidisi
    Last Modified in 02/19/21
    701029702002
    [29] † Titanochelon bolivari
    Last Modified in 02/19/21
    3513713203002
    [30] † Paleotestudo canetotiana
    Last Modified in 02/19/21
    108656101002
    [31] † Achilemys cassouleti
    Last Modified in 02/19/21
    103676500002
    [32] † Hadrianus corsoni
    Last Modified in 02/19/21
    104666301002
    [33] † Testudo eocaenica
    Last Modified in 02/19/21
    113655700002
    [34] Testudo kaiseni
    Last Modified in 02/19/21
    127424201002
    [35] † Oligopherus laticuneus
    Last Modified in 02/19/21
    91787801002
    [36] † Testudo marmorum
    Last Modified in 02/19/21
    111585801002
    [37] † Titanochelon perpiniana
    Last Modified in 02/19/21
    87807804002
    [38] Testudo promarginata
    Last Modified in 02/19/21
    111595900002
    [39] † Testudo sharanensis
    Last Modified in 02/19/21
    115545301002
    [40] † Pelorochelon soriana
    Last Modified in 02/19/21
    106706400002
    [41] † Titanochelon vitodurana
    Last Modified in 02/19/21
    81888702002
    [42] † Titanochelon schafferi
    Last Modified in 02/19/21
    101676702002


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    type number of views Individual items viewed (where applicable)
    Total project views2192
    Project overview232
    Matrix views160Data matrix (20 views); Matrix landing page (140 views);
    Media views497Media search (420 views); M739129 (77 views);
    Taxon list830
    Bibliography91
    Views for media list148
    Specimen list234




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    Total downloads from project51
    Matrix downloads7Data matrix (7 downloads);
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