Project 199: G. Giribet, P. P. Sharma, L. R. Benavides, S. L. Boyer, R. M. Clouse, B. L. de Bivort, D. Dimitrov, G. Y. Kawauchi, J. Y. Murienne, P. J. Schwendinger. 2012. Evolutionary and biogeographical history of an ancient and global group of arachnids (Arachnida: Opiliones: Cyphophthalmi) with a new taxonomic arrangement. Biological Journal of the Linnean Society. 105 (1):92-130.
Specimen: Paramiopsalis eduardoi (MCZ /DNA101878:MCZ DNA101878)
View: dorsal

Abstract

We investigate the phylogeny, biogeography, time of origin and diversification, ancestral area reconstruction and large-scale distributional patterns of an ancient group of arachnids, the harvestman suborder Cyphophthalmi. Analysis of molecular and morphological data allow us to propose a new classification system for the group; Pettalidae constitutes the infraorder Scopulophthalmi new clade, sister group to all other families, which are divided into the infraorders Sternophthalmi new clade and Boreophthalmi new clade. Sternophthalmi includes the families Troglosironidae, Ogoveidae, and Neogoveidae; Boreophthalmi includes Stylocellidae and Sironidae, the latter family of questionable monophyly. The internal resolution of each family is discussed and traced back to its geological time origin, as well as to its original landmass, using methods for estimating divergence times and ancestral area reconstruction. The origin of Cyphophthalmi can be traced back to the Carboniferous, whereas the diversification time of most families ranges between the Carboniferous and the Jurassic, with the exception of Troglosironidae, whose current diversity originates in the Cretaceous/Tertiary. Ancestral area reconstruction is ambiguous in most cases. Sternophthalmi is traced back to an ancestral land mass that contained New Caledonia and West Africa in the Permian, whereas the ancestral landmass for Neogoveidae included the south-eastern USA and West Africa, dating back to the Triassic. For Pettalidae, most results include South Africa, or a combination of South Africa with the Australian plate of New Zealand or Sri Lanka, as the most likely ancestral landmass, back in the Jurassic. Stylocellidae is reconstructed to the Thai-Malay Penisula during the Jurassic. Combination of the molecular and morphological data results in a hypothesis for all the cyphophthalmid genera, although the limited data available for some taxa represented only in the morphological partition negatively affects the phylogenetic reconstruction by decreasing nodal support in most clades. However, it resolves the position of many monotypic genera not available for molecular analysis, such as Iberosiro, Odontosiro, Speleosiro, Managotria or Marwe, although it does not place Shearogovea or Ankaratra within any existing family. The biogeographical data show a strong correlation between relatedness and formerly adjacent landmasses, and oceanic dispersal does not need to be postulated to explain disjunct distributions, especially when considering the time of divergence. The data also allow testing of the hypotheses of the supposed total submersion of New Zealand and New Caledonia, clearly falsifying submersion of the former, although the data cannot reject the latter.


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Article DOI: 10.1111/j.1095-8312.2011.01774.x

Project DOI: 10.7934/P199, http://dx.doi.org/10.7934/P199
This project containsMatrices
  • 313 Media
  • 1 Matrix
  • 1 Document
  • 318 Taxa
  • 33 Specimens
  • 76 Characters
Total size of project's media files: 1150 MB

Download Project SDD File
Total scored cells: 13939
Total media associated with cells: 51
Total labels associated with cell media: 2
Characters
Total characters: 76
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 191
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:76/0
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MorphoBank Project 199
  • Creation Date:
    08 March 2008
  • Publication Date:
    01 April 2012
  • Date of Last Publication of Partition:
    25 March 2013
  • Project views: 26459
  • Matrix downloads: 5

    This research
    supported by



    Members

    member name taxa specimens media media
    notes
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    notes  media
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    (scored, NPA, "-")
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    Gonzalo Giribet
    Project Administrator
    Last logged in 04/29/16
    2826407421009653
    (8846, 182, 625)
    95100
    Maureen Admin
    Full membership
    Last logged in 05/11/17
    000000000
    (0, 0, 0)
    0000
    Ligia R. Benavides
    Full membership
    Last logged in 04/24/17
    66401000516
    (484, 1, 31)
    0000
    Sarah Boyer
    Full membership
    Last logged in 04/11/11
    01000000344
    (337, 0, 7)
    0000
    Ronald Clouse
    Full membership
    Last logged in 05/18/12
    2000000001823
    (1733, 0, 90)
    0000
    MorphoBank Curator
    Full membership
    Last logged in 05/05/17
    100000000
    (0, 0, 0)
    0000
    Benjamin de Bivort
    Full membership
    Last logged in 03/07/11
    80001100917
    (882, 0, 35)
    16000
    Seth Kaufman
    Full membership
    Last logged in 05/22/16
    010000000
    (0, 0, 0)
    0000
    Maria Passarotti
    Last logged in 03/15/18
    000000000
    (0, 0, 0)
    0000
    Prashant P. Sharma
    Full membership
    Last logged in 04/13/13
    00000000686
    (662, 0, 24)
    0000


    Taxonomic Overview for Matrix 'M561' (271 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] 'Siro' beschkovi
    Last Modified in 12/05/13
    58151503009
    [2] Cyphophthalmus radjai
    Last Modified in 02/21/14
    6666604009
    [3] Cyphophthalmus gjorgjevici
    Last Modified in 12/05/13
    28454503009
    [4] Cyphophthalmus hlavaci
    Last Modified in 12/05/13
    37363603009
    [5] Cyphophthalmus corfuanus
    Last Modified in 12/05/13
    37353504009
    [6] Cyphophthalmus duricorius
    Last Modified in 12/05/13
    8636305009
    [7] Cyphophthalmus eratoae
    Last Modified in 12/05/13
    58151503009
    [8] Cyphophthalmus ere
    Last Modified in 12/05/13
    33404003009
    [9] Cyphophthalmus gordani
    Last Modified in 12/05/13
    38353503009
    [10] Cyphophthalmus markoi
    Last Modified in 12/05/13
    36373703009
    [11] Cyphophthalmus martensi
    Last Modified in 12/05/13
    30424204009
    [12] Cyphophthalmus minutus
    Last Modified in 12/05/13
    38353503009
    [13] Cyphophthalmus montenegrinus
    Last Modified in 12/05/13
    687701009
    [14] 'Siro' carpaticus
    Last Modified in 12/05/13
    58151503009
    [15] Cyphophthalmus noctiphilus
    Last Modified in 12/05/13
    687701009
    [16] Cyphophthalmus ognjenovici
    Last Modified in 12/05/13
    33393904009
    [17] Cyphophthalmus ohridanus
    Last Modified in 12/05/13
    687701009
    [18] Cyphophthalmus paradoxus
    Last Modified in 12/05/13
    18545404009
    [19] Cyphophthalmus rumijae
    Last Modified in 12/05/13
    36373703009
    [20] Cyphophthalmus serbicus
    Last Modified in 12/05/13
    10616105009
    [21] Cyphophthalmus silhavyi
    Last Modified in 12/05/13
    687701009
    [22] Cyphophthalmus teyrovskyi
    Last Modified in 12/05/13
    39343403009
    [23] Cyphophthalmus trebinjanus
    Last Modified in 12/05/13
    21515104009
    [24] Cyphophthalmus zetae
    Last Modified in 12/05/13
    38353503009
    [25] Iberosiro distylos
    Last Modified in 12/05/13
    9585845009
    [26] Odontosiro lusitanicus
    Last Modified in 12/05/13
    18535205009
    [27] Paramiopsalis ramblae
    Last Modified in 10/30/16
    49272700009
    [28] Paramiopsalis eduardoi
    Last Modified in 12/05/13
    10636303009
    [29] Paramiopsalis ramulosus
    Last Modified in 12/05/13
    76622034509
    [30] Paramiopsalis anadonae
    Last Modified in 10/30/16
    19535313009
    [31] Parasiro coiffaiti
    Last Modified in 12/05/13
    7656404109
    [32] Parasiro corsicus
    Last Modified in 12/05/13
    15585803009
    [33] Parasiro minor
    Last Modified in 12/05/13
    9636304009
    [34] Siro acaroides
    Last Modified in 12/05/13
    7656504009
    [35] Siro boyerae
    Last Modified in 12/05/13
    4686804009
    [36] Siro calaveras
    Last Modified in 12/05/13
    9606034009
    [37] Siro clousi
    Last Modified in 12/05/13
    60151501009
    [38] Siro crassus
    Last Modified in 12/05/13
    64111101009
    [39] Siro exilis
    Last Modified in 12/05/13
    9636304009
    [40] Siro kamiakensis
    Last Modified in 12/05/13
    9636304009
    [41] Siro ligiae
    Last Modified in 12/05/13
    64111101009
    [42] † Siro platypedibus
    Last Modified in 12/05/13
    62121202009
    [43] Siro rubens
    Last Modified in 12/05/13
    7656504009
    [44] Siro shasta
    Last Modified in 12/05/13
    6656214309
    [45] Siro sonoma
    Last Modified in 12/05/13
    18545404009
    [46] Siro valleorum
    Last Modified in 12/05/13
    6666604009
    [47] Suzukielus sauteri
    Last Modified in 12/05/13
    7666603009
    [48] Ankaratra franzi
    Last Modified in 12/05/13
    27434324009
    [49] Aoraki calcarobtusa calcarobtusa
    Last Modified in 12/05/13
    16595901009
    [50] Aoraki calcarobtusa westlandica
    Last Modified in 12/05/13
    8656503009
    [51] Aoraki crypta
    Last Modified in 12/05/13
    6686802009
    [52] Aoraki denticulata denticulata
    Last Modified in 12/05/13
    6686802009
    [53] Aoraki denticulata major
    Last Modified in 12/05/13
    8666602009
    [54] Aoraki granulosa
    Last Modified in 12/05/13
    9646403009
    [55] Aoraki healyi
    Last Modified in 12/05/13
    7676702009
    [56] Aoraki inerma inerma
    Last Modified in 12/05/13
    6686802009
    [57] Aoraki longitarsa
    Last Modified in 12/05/13
    8666602009
    [58] Aoraki sp_dna101126
    Last Modified in 12/05/13
    9646403009
    [59] Aoraki tumidata
    Last Modified in 12/05/13
    8656503009
    [60] Austropurcellia arcticosa
    Last Modified in 12/05/13
    12626202009
    [61] Austropurcellia capricornia
    Last Modified in 12/05/13
    18575701009
    [62] Austropurcellia daviesae
    Last Modified in 12/05/13
    8666602009
    [63] Austropurcellia forsteri
    Last Modified in 12/05/13
    9656502009
    [64] Austropurcellia scoparia
    Last Modified in 12/05/13
    6686802009
    [65] Austropurcellia woodwardi
    Last Modified in 12/05/13
    47292900009
    [66] Chileogovea jocasta
    Last Modified in 12/05/13
    14595903009
    [67] Chileogovea oedipus
    Last Modified in 12/05/13
    9626205009
    [68] Chileogovea sp_dna100490
    Last Modified in 12/05/13
    15565605009
    [69] Karripurcellia harveyi
    Last Modified in 12/05/13
    12595905009
    [70] Karripurcellia peckorum
    Last Modified in 12/05/13
    9626205009
    [71] Karripurcellia sierwaldae
    Last Modified in 12/05/13
    12595905009
    [72] Manangotria taolanaro
    Last Modified in 12/05/13
    25434326009
    [73] Neopurcellia salmoni
    Last Modified in 12/05/13
    18545404009
    [74] Parapurcellia fissa
    Last Modified in 12/05/13
    24514701009
    [75] Parapurcellia monticola
    Last Modified in 12/05/13
    12615703009
    [76] Parapurcellia rumpiana
    Last Modified in 12/05/13
    18565402009
    [77] Parapurcellia silvicola
    Last Modified in 12/05/13
    12615903009
    [78] Parapurcellia amatolae
    Last Modified in 12/05/13
    13616102009
    [79] Parapurcellia convexa
    Last Modified in 12/05/13
    12626202009
    [80] Parapurcellia minuta
    Last Modified in 12/05/13
    12616103009
    [81] Parapurcellia natalia
    Last Modified in 12/05/13
    14606002009
    [82] Parapurcellia staregai
    Last Modified in 12/05/13
    14606002009
    [83] Pettalus brevicauda
    Last Modified in 12/05/13
    13595904009
    [84] Pettalus cimiciformis
    Last Modified in 12/05/13
    15575704009
    [85] Pettalus lampetides
    Last Modified in 12/05/13
    8646404009
    [86] Pettalus sp_dna101288
    Last Modified in 12/05/13
    21505005009
    [87] Pettalus sp_dna101285
    Last Modified in 12/05/13
    12595905009
    [88] Pettalus sp_dna101286
    Last Modified in 12/05/13
    11606005009
    [89] Pettalus sp_dna101282
    Last Modified in 12/05/13
    11606005009
    [90] Pettalus sp_dna101283
    Last Modified in 12/05/13
    12595905009
    [91] Pettalus sp_dna101287
    Last Modified in 12/05/13
    13585805009
    [92] Pettalus thwaitesi
    Last Modified in 12/05/13
    7646405009
    [93] Purcellia illustrans
    Last Modified in 12/05/13
    7666503009
    [94] Parapurcellia peregrinator
    Last Modified in 12/05/13
    16585802009
    [95] Purcellia transvaalica
    Last Modified in 12/05/13
    16585702009
    [96] Purcellia griswoldi
    Last Modified in 12/05/13
    9656502009
    [97] Purcellia lawrencei
    Last Modified in 12/05/13
    13616102009
    [98] Rakaia 134 431
    Last Modified in 12/05/13
    19545403009
    [99] Rakaia sp_dna101297
    Last Modified in 12/05/13
    13606003009
    [100] Rakaia antipodiana
    Last Modified in 12/05/13
    14606002009
    [101] Rakaia dorothea
    Last Modified in 12/05/13
    14606002009
    [102] Rakaia florensis
    Last Modified in 12/05/13
    14606002009
    [103] Rakaia isolata
    Last Modified in 12/05/13
    741101009
    [104] Rakaia sp_dna100958
    Last Modified in 12/05/13
    14606002009
    [105] Rakaia sp_dna101293
    Last Modified in 12/05/13
    15595902009
    [106] Rakaia lindsayi
    Last Modified in 12/05/13
    13616102009
    [107] Rakaia macra
    Last Modified in 12/05/13
    11636302009
    [108] Rakaia magna australis
    Last Modified in 12/05/13
    13616102009
    [109] Rakaia media
    Last Modified in 12/05/13
    13616102009
    [110] Rakaia minutissima
    Last Modified in 12/05/13
    14606002009
    [111] Rakaia pauli
    Last Modified in 12/05/13
    14606002009
    [112] Rakaia solitaria
    Last Modified in 12/05/13
    13616102009
    [113] Rakaia sorenseni digitata
    Last Modified in 12/05/13
    13606003009
    [114] Rakaia sorenseni sorenseni
    Last Modified in 12/05/13
    14606002009
    [115] Rakaia sp_dna101807
    Last Modified in 12/05/13
    14595903009
    [116] Rakaia stewartiensis
    Last Modified in 12/05/13
    14606002009
    [117] Rakaia uniloca
    Last Modified in 12/05/13
    14595903009
    [118] Rakaia sp_dna100954
    Last Modified in 12/05/13
    12626202009
    [119] Speleosiro argasiformis
    Last Modified in 12/05/13
    9646403009
    [120] Marwe coarctata
    Last Modified in 12/05/13
    11585807009
    [121] Shearogovea mexasca
    Last Modified in 12/05/13
    9585827009
    [122] Metasiro americanus
    Last Modified in 12/05/13
    1717104009
    [123] Brasiliogovea yacambui
    Last Modified in 04/15/17
    704402009
    [124] Brasilogovea microphaga
    Last Modified in 12/05/13
    2676716009
    [125] Brasilogovea chiribiqueta
    Last Modified in 12/05/13
    3686805009
    [126] Canga renatae
    Last Modified in 12/05/13
    8626215009
    [127] Huitaca ventralis
    Last Modified in 12/05/13
    2686715009
    [128] Huitaca sp tama_dna101671
    Last Modified in 12/05/13
    4676705009
    [129] Huitaca sp boyaca_dna101407
    Last Modified in 12/05/13
    3686805009
    [130] Huitaca sp caldas_dna101681
    Last Modified in 12/05/13
    2696905009
    [131] Huitaca sp bitaco_dna101683
    Last Modified in 12/05/13
    5666605009
    [132] Huitaca sp depressa_dna102150
    Last Modified in 12/05/13
    3686805009
    [133] Huitaca sp _dna104646
    Last Modified in 12/05/13
    3686805009
    [134] Huitaca sp._dna105823
    Last Modified in 12/05/13
    48252503009
    [135] Leggogovia pabsgarnoni
    Last Modified in 04/15/17
    122929341009
    [136] Metagovea disparunguis
    Last Modified in 12/05/13
    34393903009
    [137] Metagovea sp nambi_dna101410
    Last Modified in 12/05/13
    5676704009
    [138] Metagovea planada
    Last Modified in 04/15/17
    6666604009
    [139] Metagovea philipi
    Last Modified in 12/05/13
    4686804009
    [140] Metagovea sp caldas_dna101680
    Last Modified in 12/05/13
    5676704009
    [141] Metagovea sp planada_dna101654
    Last Modified in 12/05/13
    5676704009
    [142] Metagovea sp valle_dna101685
    Last Modified in 12/05/13
    5676704009
    [143] Metagovea sp tatama_dna102151m
    Last Modified in 12/05/13
    6666604009
    [144] Metagovea sp leticia_dna101408
    Last Modified in 12/05/13
    5676704009
    [145] Metagovea sp chiriboga
    Last Modified in 12/05/13
    55202001009
    [146] Metagovea sp huila_dna101670
    Last Modified in 12/05/13
    7656504009
    [147] Metagovea sp nambi_dna101409
    Last Modified in 12/05/13
    4686804009
    [148] Metagovea sp pasto_dna101642
    Last Modified in 12/05/13
    4686804009
    [149] Metagovea sp valle_dna101686
    Last Modified in 12/05/13
    4686804009
    [150] Microgovia oviformis
    Last Modified in 04/15/17
    8646404009
    [151] Microgovia ayanganna
    Last Modified in 04/15/17
    7646405009
    [152] Neogovea branstetteri_52990
    Last Modified in 04/15/17
    9626205009
    [153] Neogovea enigmatica
    Last Modified in 04/16/17
    103938271009
    [154] Neogovea immsi
    Last Modified in 12/05/13
    16555505009
    [155] Neogovea kamakusa
    Last Modified in 12/05/13
    17555504009
    [156] Neogovea kartabo
    Last Modified in 12/05/13
    21515104009
    [157] Neogovea virginie
    Last Modified in 12/05/13
    10595916009
    [158] Parogovia sironoides
    Last Modified in 12/05/13
    1717104009
    [159] Parogovia cf_sironoides
    Last Modified in 12/05/13
    1717104009
    [160] Parogovia gabonica
    Last Modified in 12/05/13
    10626204009
    [161] Parogovia pabsgarnoni
    Last Modified in 12/05/13
    12606004009
    [162] Parogovia montealensis
    Last Modified in 04/15/17
    15575704009
    [163] Parogovia sp campo2_dna104618
    Last Modified in 12/05/13
    252020301009
    [164] Parogovia sp koupe_dna104615
    Last Modified in 12/05/13
    17555504009
    [165] Parogovia prietoi
    Last Modified in 12/05/13
    3686805009
    [166] Parogovia sp_dna101058
    Last Modified in 12/05/13
    60151501009
    [167] Metagovea sp apaporis
    Last Modified in 12/05/13
    52222202009
    [168] Metagovea sp chiribiquete dna101661
    Last Modified in 12/05/13
    62121202009
    [169] Metagovea sp buga (florezi)
    Last Modified in 12/05/13
    21525203009
    [170] Metagovea sp guacharos_dna101660
    Last Modified in 12/05/13
    16575703009
    [171] Metagovea sp_dna101678 small
    Last Modified in 12/05/13
    48252503009
    [172] Metagovea sp heremberg
    Last Modified in 12/05/13
    62131301009
    [173] Metagovea sp tena
    Last Modified in 12/05/13
    52222202009
    [174] Metagovea dna101667 (picachos)
    Last Modified in 12/05/13
    54202002009
    [175] Metagovea dna101677 (big)
    Last Modified in 12/05/13
    20535303009
    [176] Metagovea sp yanayacu
    Last Modified in 12/05/13
    63121201009
    [177] Neogovea sp cayenne
    Last Modified in 12/05/13
    53212102009
    [178] Neogovea sp purace
    Last Modified in 12/05/13
    62131301009
    [179] Neogovea sp trinidad (sp 7)
    Last Modified in 12/05/13
    62131301009
    [180] Neogovea sp suriname
    Last Modified in 12/05/13
    52222202009
    [181] Neogovea sp arima (sp 8)
    Last Modified in 12/05/13
    63121201009
    [182] Neogovea turumai
    Last Modified in 12/05/13
    30444402009
    [183] Waiwaigovia schultzi
    Last Modified in 04/15/17
    15575704009
    [184] Troglosiro aelleni
    Last Modified in 12/05/13
    7656504009
    [185] Troglosiro longifossa
    Last Modified in 12/05/13
    3696904009
    [186] Troglosiro brevifossa
    Last Modified in 12/05/13
    13606003009
    [187] Troglosiro juberthiei
    Last Modified in 12/05/13
    6666604009
    [188] Troglosiro monteithi
    Last Modified in 12/05/13
    8646404009
    [189] Troglosiro ninqua
    Last Modified in 12/05/13
    8646404009
    [190] Troglosiro oscitatio
    Last Modified in 12/05/13
    11626203009
    [191] Troglosiro platnicki
    Last Modified in 12/05/13
    11626203009
    [192] Troglosiro raveni
    Last Modified in 12/05/13
    11626203009
    [193] Troglosiro sheari
    Last Modified in 12/05/13
    12616103009
    [194] Troglosiro tillierorum
    Last Modified in 12/05/13
    10636303009
    [195] Troglosiro urbanus
    Last Modified in 12/05/13
    8646404009
    [196] Troglosiro wilsoni
    Last Modified in 12/05/13
    8646404009
    [197] Ogovea cameroonensis
    Last Modified in 12/05/13
    8626215009
    [198] Ogovea grossa
    Last Modified in 12/05/13
    152525342009
    [199] Ogovea nasuta
    Last Modified in 12/05/13
    11575735009
    [200] Fangensis cavernarum
    Last Modified in 12/05/13
    10646402009
    [201] Fangensis insulanus
    Last Modified in 12/05/13
    10636303009
    [202] Fangensis leclerci
    Last Modified in 12/05/13
    23515102009
    [203] Fangensis spelaeus
    Last Modified in 12/05/13
    8656503009
    [204] Leptopsalis beccarii
    Last Modified in 12/05/13
    10606006009
    [205] Leptopsalis lydekkeri_dna101064
    Last Modified in 12/05/13
    15565605009
    [206] Leptopsalis novaguinea_dna101510
    Last Modified in 12/05/13
    17545405009
    [207] Leptopsalis sp_dna101093_496
    Last Modified in 12/05/13
    15565605009
    [208] Leptopsalis sp_dna101474
    Last Modified in 12/05/13
    18545404009
    [209] Leptopsalis sp_dna101478
    Last Modified in 12/05/13
    19535304009
    [210] Leptopsalis sp_dna101483
    Last Modified in 12/05/13
    14575705009
    [211] Leptopsalis sp_dna101489
    Last Modified in 12/05/13
    14575705009
    [212] Leptopsalis sp_dna101500
    Last Modified in 12/05/13
    733300009
    [213] Leptopsalis sp_dna101514
    Last Modified in 12/05/13
    14585804009
    [214] Leptopsalis sp_dna101929
    Last Modified in 12/05/13
    733300009
    [215] Leptopsalis sp_dna101930
    Last Modified in 12/05/13
    15565605009
    [216] Leptopsalis sp_dna101931
    Last Modified in 12/05/13
    733300009
    [217] Leptopsalis sp_dna101932
    Last Modified in 12/05/13
    26464604009
    [218] Leptopsalis sp_dna101934
    Last Modified in 12/05/13
    733300009
    [219] Leptopsalis sp_dna101935
    Last Modified in 12/05/13
    733300009
    [220] Leptopsalis sp_dna101936
    Last Modified in 12/05/13
    733300009
    [221] Leptopsalis sp_dna101937
    Last Modified in 12/05/13
    17555504009
    [222] Leptopsalis sp_dna101938
    Last Modified in 12/05/13
    14575705009
    [223] Leptopsalis sp_dna101942
    Last Modified in 12/05/13
    733300009
    [224] Leptopsalis sp_dna101944_5
    Last Modified in 12/05/13
    24484804009
    [225] Leptopsalis sp_dna102033_48
    Last Modified in 12/05/13
    26464604009
    [226] Leptopsalis sp_dna102039
    Last Modified in 12/05/13
    27454504009
    [227] Leptopsalis sp_dna102042
    Last Modified in 12/05/13
    26464604009
    [228] Leptopsalis sp_dna102061
    Last Modified in 12/05/13
    27454504009
    [229] Leptopsalis sp_dna103250
    Last Modified in 12/05/13
    26464604009
    [230] Leptopsalis sp_dna102032
    Last Modified in 12/05/13
    741101009
    [231] Meghalaya annandalei
    Last Modified in 12/05/13
    11585825009
    [232] Meghalaya sp_dna101094_500
    Last Modified in 12/05/13
    18545404009
    [233] Meghalaya sp_dna101494_506_765
    Last Modified in 12/05/13
    29444403009
    [234] Megahalaya sp_dna101767
    Last Modified in 12/05/13
    27464603009
    [235] Meghalaya sp_dna102051
    Last Modified in 12/05/13
    27464603009
    [236] Meghalaya sp_dna103242_3_4
    Last Modified in 12/05/13
    26474703009
    [237] Meghalaya sp_dna103251
    Last Modified in 12/05/13
    25494902009
    [238] Meghalaya sp_dna103265
    Last Modified in 12/05/13
    28464602009
    [239] Miopsalis pulicaria
    Last Modified in 12/05/13
    113333311009
    [240] Miopsalis sp_dna101468_950
    Last Modified in 12/05/13
    26484802009
    [241] Miopsalis sp_dna101517
    Last Modified in 12/05/13
    16575703009
    [242] Miopsalis sp_dna102032_53_58
    Last Modified in 12/05/13
    16565604009
    [243] Miopsalis sp_dna103249
    Last Modified in 12/05/13
    31424203009
    [244] Miopsalis sp_dna103254
    Last Modified in 12/05/13
    28464602009
    [245] Miopsalis sp_dna101519
    Last Modified in 12/05/13
    59161601009
    [246] Miopsalis sp_dna103259
    Last Modified in 12/05/13
    64111101009
    [247] Stylocellus collinsi
    Last Modified in 12/05/13
    26474703009
    [248] Stylocellus dumoga
    Last Modified in 12/05/13
    21505005009
    [249] Stylocellus globosus
    Last Modified in 12/05/13
    15575704009
    [250] Stylocellus gryllospecus
    Last Modified in 12/05/13
    12616103009
    [251] Stylocellus hillyardi
    Last Modified in 12/05/13
    16565604009
    [252] Stylocellus javanus
    Last Modified in 12/05/13
    715500009
    [253] Stylocellus kinabalu
    Last Modified in 12/05/13
    715500009
    [254] Stylocellus laevichelis
    Last Modified in 12/05/13
    715500009
    [255] Stylocellus leakeyi
    Last Modified in 12/05/13
    715500009
    [256] Stylocellus lionotus
    Last Modified in 12/05/13
    715500009
    [257] Stylocellus modestus
    Last Modified in 12/05/13
    715500009
    [258] Stylocellus mulu
    Last Modified in 12/05/13
    715500009
    [259] Stylocellus pangrango
    Last Modified in 12/05/13
    715500009
    [260] Stylocellus pocockii
    Last Modified in 12/05/13
    715500009
    [261] Stylocellus ramblae
    Last Modified in 12/05/13
    15565405209
    [262] Stylocellus sabah
    Last Modified in 12/05/13
    27464603009
    [263] Stylocellus silhavyi
    Last Modified in 12/05/13
    724400009
    [264] Stylocellus spinifrons
    Last Modified in 12/05/13
    715500009
    [265] Stylocellus sp. irian jaya
    Last Modified in 12/05/13
    58161602009
    [266] Stylocellus sulcatus
    Last Modified in 12/05/13
    715500009
    [267] Stylocellus sumatranus
    Last Modified in 12/05/13
    17535324009
    [268] Stylocellus tarumpitao
    Last Modified in 12/05/13
    715500009
    [269] Stylocellus tambusisi
    Last Modified in 12/05/13
    17545405009
    [270] Stylocellus thorellii
    Last Modified in 12/05/13
    715500009
    [271] Stylocellus weberii
    Last Modified in 12/05/13
    715500009


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    Project overview686
    Documents list244
    Matrix views359Matrix landing page (314 views); Morphological data matrix of the Opiliones suborder Cyphophthalmi (Arthropoda: Arachnida) (45 views);
    Bibliography1694
    Taxon list2014
    Specimen list2969
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    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project40
    Project downloads35
    Matrix downloads5Morphological data matrix of the Opiliones suborder Cyphophthalmi (Arthropoda: Arachnida) (5 downloads);